S&W made a change to the sear deactivation lever early in the M&P's production. This was reportedly because some LE users couldn't remember to keep the lever in the down position during reassembly, when they were trying to install the slide.
If the early lever design was in the up position as the slide was being installed, the slide's pickup rail could catch against the lever, risking damage to it. Oops. User error.
S&W engineers revised the lever so it would clear the pickup rail under those conditions.
In this picture the old style sear release lever is on the left, and the new style lever is on the right. (The left block also has the mag safety lever in it, but no ejector, BTW.) The first armorer manual (Feb '06) shows the original lever, while the first revision (Aug '08) shows both types of levers in different pictures.