Simply firing the gun makes those marks. On discharge, the gun recoils backward, and the cylinder, attached to the crane through it's inertia, slams forward into the frame. It tends to bounce a bit during cycling, and the ratchet hub impacts the rear of the frame.
That's why excessive cylinder fore-and-aft movement (endplay) increases as a gun is fired more, until headspace increases and the barrel-to-cylinder gap decreases to the point of unserviceability. The hotter the loads, the lighter the gun, the more pronounced the movement is. Uncorrected excessive endplay can be very damaging to a revolver.
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