Google the Houston Warehouse experiment and reread it annually.
How many shooters here have a custom made rifle in 22 PPC shortened .05” where the barrel has been lapped to the receiver for 100% thread contact?
How many here have their triggers set to 1/4 of an once?
How many here go to the extensive case prep that was in this article?
How many here jam their bullets into the rifling?
Bottom line, too many people think if they copy what benchrest shooter do with their custom rifles with tight neck chambers it will help their off the shelf factory rifles.
The OP question was "What’s the significance of using bushing dies?"
Answer, spending much more money on bushing dies that will have more neck runout than a standard full length die.
When reducing the neck diameter .004 or more with a bushing die it is recommended to reduce the neck diameter in two steps to help reduce neck runout.
Now ask yourself "WHY" so many reloaders prefer the Lee collet die over bushing dies.
Answer, they get "LESS" bullet runout with a Lee collet die vs a bushing die.
With a bushing die and not use the expander you will need to neck turn your brass.
If you do not neck turn and use the expander Redding recommends sizing the neck .004 smaller than bullet diameter. Meaning you are still working the cases neck.
Below is a Forster full length benchrest die with its high mounted floating expander. And these Forster dies produce the most concentric resized cases of any die I have ever owned. The neck of the case is held and centered in the neck of the die when the expander enters the case neck. And this prevents the expander from pulling the case neck off center and inducing case neck runout.
Below is the Forster benchrest seating die that holds the case and bullet in perfect alignment during bullet seating. And standard seating dies do not do this and can allow the bullet to tilt during seating.
The Forster die is much cheaper than a bushing die and their benchrest seating die is cheaper the Reddings. And with both of these dies you will reload cartridges with little to no bullet runout.
"What’s the significance of using bushing dies?"
Spending more money for a bushing die in a off the shelf factory rifle that you didn't need to spend.