I think "Harry" covered the bases fairly well.
I went through the SW99/P99 armorer class (taught by S&W) 3 times. I carried an issued one for a few years, and I own 2 of them. I've put some tens of thousands of rounds through various examples of them and helped maintain more than 50 agency-owned examples, and a number of other folk's personally-owned 99's.
S&W machined and through-hardened their own slides and barrels for the SW99/99NJ/990L's. They had them sent out for Melonite QP surface hardening (versus the Tenifer QPQ used by Walther, since S&W used stainless steel for their slides and barrels). Everything else came from Walther.
S&W did request some minor design changes to the frame, a couple of which I was told Walther later adopted in their own guns. Non-hooked frame tang above the web of the hand, different standardized accessory rail, exposed toe of magazine base to remove a stuck mag ... to name a couple off the top of my head.
We were told that it was S&W engineers, using their high speed imaging, that eventually identified the cause of the erratic premature slide-lock (rounds remaining in mag) that had been reported with early P99 .40's and SW99 .40's. It involved the magazine shape at the mag catch cutout, and after S&W gave the info to Walther, Walther and Mec-Gar revised the magazine bodies and followers.
S&W also changed a couple of the springs used in the SW99/990L's,like using an optional extractor spring (and reversing its installed orientation), and the slide stop lever spring (hooked end instead of closed loop end).
I saw some ongoing barrel design revisions in the SW99's, including the chamber mouth shape and rollover point, as well as the clearance cuts on the bottom of the barrel to better clear the top of the recoil spring assembly's coils.
Good guns. I've seen them withstand extended use and abuse.
Gotta run. Just some quick thoughts.